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Odum Conference Poster Abstracts

Theme A     Theme B     Theme C     Theme D     Poster Home    Printer-friendly Abstracts

Theme B: Basic research on dynamic aspects of invasions

* indicates presenting author.

Adams, David*; Roblee, Kenneth; Stone, Ward – NYS DEC Office of Invasive Species Coordination

NYS waterbird mortality as a result of Type E botulism and nonnative invasive species

Periodic waterbird mortality, due to type E botulism (Clostridium botulinum) intoxication, has been documented on the Great Lakes since the 1960s. The most recent series of events are unique in that they appear to be linked to population increases of two nonnative invasive species, the quagga mussel (Dreissena bugensis) and the round goby (Neogobius melanostomus).

The first documented mortality in New York State occurred in the eastern basin of Lake Erie November 2000. To monitor impacts on waterbirds, thirteen 500-meter shoreline transects were established and monitored weekly. Surveys were repeated during fall 2001 through 2008. By extrapolation we estimate that up to 5415, 2862, 17301, 3008, 5943, 2297, 4375, 8915 and 2005 waterbirds may have died.

In an effort to detect increases in the geographic extent of these annual mortality events, forty-seven 500-meter transects were placed at five mile intervals along the Lake Ontario shoreline during 2002. These transects were monitored weekly during the peak of Common Loon (Gavia immer) migration 2002-2008. By extrapolation we estimate that up to 1046, 1529, 1693, 1193, 4933, 3650 and 1628 waterbirds may have perished.

The waterbird species with the greatest mortality differed for the 2000-02 mortality events. Redbreasted Merganser (Mergus serrator) had a predicted mortality of 2479 in 2000, Common Loon 1149 during 2001, and Long-tailed Duck (Clangula hyemalis) 13219 in 2002. Common Loon experienced the greatest mortality during the 2003-8 events, ranging from 1656 to 10040.

Allen, Jenica M*1; Ibáñez, Inés2, Wilson, Adam M1; Treanor, Sarah A1; Silander, John A1 – (1) University of Connecticut; (2) University of Michigan

Identifying hot spots of plant species invasions and assessing foci of further spread

Invasive species are a global ecological and economic issue, making inference on where to target early control and eradication efforts critical. We aimed to forecast hotspots of invasion in terms of invasive species richness and high abundance of particular species by combining climate, land-use/land-cover (LULC), and habitat information in spatially-explicit hierarchical Bayesian models. Species occurrence and abundance data were derived from the Invasive Plant Atlas of New England (IPANE), demonstrating the potential for fruitful collaborations between researchers, citizen scientists, and resource managers. Species–specific analyses included prominent plant invaders in New England: Berberis thunbergii (Japanese barberry), Celastrus orbiculatus (oriental bittersweet), Elaeagnus umbellata (autumn olive), Euonymus alata (burning bush), and Rosa multiflora (multiflora rose). Projections were made using two future climate models (CCSM3 and UKMO-HadCM3) in addition to current climate.

Both the richness and abundance models indicate that climate variables were most influential in determining the incidence or cover of invasives, followed by site and LULC variables. Invasive species richness was higher in warmer areas with lower annual and highly seasonally variable precipitation. Edge habitats increased and evergreen forest and cropland LULC decreased invasive richness. The abundance analysis showed differences in the direction and magnitude of the effect of each predictive variable among the five species, demonstrating the need for both synthetic and species-specific approaches to prediction and management. Model predictions suggest that all species, except E.umbellata, may be more abundant relative to current climate under moderate predictions (CCSM3), but that their incidence may be reduced under extreme climate predictions (HadCM3).

Allstadt, Andrew*; O’Malley, L; Kozma, B; Korniss, G; Racz, Z; Caraco, T; Newman, J – University at Albany

Velocity and roughening of an invasive front

After establishment, many exotic species expand into the surrounding environment through local propagation, spreading as a discernable invasive front. We model this spatial advance as an individual-based, stochastic process in two dimensions, where an invasive and a resident species compete preemptively for resources. Initially, the species are separated by a linear front; the invader’s subsequent advance is governed by local dispersal and mortality events in both species. At early times, correlations in the position of leading invasive individuals develop over short distances. As the front continues to advance, the length of these correlations grows until they extend over the entire front. Velocity of the front and variance about the average front position grow with correlation length, and reach their asymptotic values as the correlation length spans the front. Consequently both asymptotic velocity and front variance exhibit specific scaling relationships with the front length, regardless of the detailed propagation and mortality processes. These relationships also suggest this ecological model belongs to the Kardar-Parisi-Zhang (KPZ) universality class; a set of spatial-growth models sharing common scaling properties. Other KPZ systems include the advance of wildfires, spatial epidemic spread, and the Eden model of tumor growth. Given that ecological invasion with local dispersal belongs to this universality class, we can specify the probability distribution of the most advanced invader, the “front runner.” We shall test hypotheses related to the length of an invasive front with a field experiment, using grass and clover as a model system.

Averill, Kristine M*1; DiTommaso, Antonio1; Mohler, Charles L1; Milbrath, Lindsey R2 – (1) Cornell University; (2) USDA-ARS Robert W. Holley Center for Agriculture and Health

Generating biological data on the invasive swallow-worts (Vincetoxicum spp.) to better inform their management using biocontrol

Pale and black swallow-wort (Vincetoxicum rossicum and V. nigrum, respectively) are nonnative, perennial, herbaceous vines in the Apocynaceae. The species are becoming increasingly problematic in the northeastern United States and southeastern Canada and management of the species has been challenging. A classical biological control program was initiated in 2004 by the USDA-ARS. Success of this biological control effort depends on the availability of plant demographic data, which can be modeled to determine which swallow-wort life stage(s) are likely to be most susceptible to control efforts. Since little is known about the establishment phase of swallow-wort’s life cycle, we conducted a field experiment in central New York State to determine establishment and survival during the first two years after sowing.

We sowed pale swallow-wort seeds in two old-fields subjected to four disturbance regimens. We hypothesized that establishment and survival would be greater in treatments with greater disturbance. At the better-drained site, overall swallow-wort establishment was 15(1)% and did not differ between treatments. At the poorly-drained site, establishment varied by treatment; mowed and control plots had greater swallow-wort establishment (10(2)%) than herbicide+tillage and herbicide plots (1.6(0.5)%). Of emerging seedlings, survival was high at both sites (70–84%). At both sites, swallow-wort total (above- and below-ground) biomass was greater in herbicide + tillage and herbicide plots than in mowed and control plots. These results indicate that pale swallow-wort establishment and survival is successful across a range of disturbance regimens and that growth is greater in more disturbed regimens. Biocontrol implications will be discussed.

Chang, Cynthia*; Smith, Melinda – Yale University

Different levels of diversity have varying impacts on invasion resistance and productivity of tallgrass prairie

Native community diversity has historically been thought to play a central role in invasion resistance by increasing interspecific trait variation. Trait variation also occurs at the intraspecific level, where phenotypic and/or genotypic diversity could confer invasion resistance via the same mechanisms. Genotypic variation within a dominant species may account for overall phenotypic trait variation within a given community, and thus may play a central role in determining invasion resistance. We conducted our study in 66 1-m2 plots located in intact tallgrass prairie. For each plot, we measured the genotypic and corresponding phenotypic variation of dominant C4 grass individuals (Andropogon gerardii) and plant community diversity. Establishment and persistence of added invasive grass seeds (A. bladhii) was monitored. Finally, we measured resource (nitrogen, water and light) availability. Multiple regression analyses found early-season soil moisture content and end of season individual biomass of the dominant species to be significantly related to invasion success (both early and late season). Genotype variation of A. gerardii was also significantly related to late season persistence of the invasive grass. This suggests that both resource availability and genotypic variation within a dominant species are important drivers in invasion resistance. Furthermore, genotypic and phenotypic variation are positively correlated which suggests that phenotypic trait variation in a dominant species could be one key mechanism driving invasion resistance. Finally, community diversity was not found to be significantly related to invasion, suggesting that it is population level diversity of the dominant species that plays an important role in tallgrass prairie under invasion.

Cohen, Jillian S*1; Maerz, John C2; Blossey, Bernd1 – (1) Cornell University; (2) University of Georgia

It’s not where you're from, it’s who you are that counts: Plant traits determine impacts on larval amphibians

Plant detritus is the primary energy source in freshwater benthic food webs. Theory suggests that plant invasions can affect consumers by changing the quality of the detrital pool, but empirical evidence for this is lacking. Tadpoles, a dominant wetland consumer, may be particularly responsive to shifts in litter quality, as they feed largely on detrital biofilms. I conducted two experiments to evaluate the impacts of invasive plant litter on tadpoles. For each experiment, I hypothesized that litter quality, including C:N, C:P, N:P, and phenolic content, would determine tadpole productivity. First, I conducted a field experiment involving four native and five invasive plant species at five wetland complexes in central New York. I installed field cages in sites dominated by a single species of each plant and added three tadpole species according to their natural phenology. I monitored cages regularly, collecting and weighing all metamorphosed individuals as they became available. Importantly, my data shows that tadpole performance did not differ in habitats dominated by native and invasive plants. Rather, my findings suggest that plant traits, irrespective of plant origin, determine amphibian performance. For my second experiment, I raised tadpoles in mesocosms containing litter from 15 populations of a single invasive species that varied widely in litter quality. Similar to my first experiment, litter traits explained much of the observed variation in tadpole productivity. These experiments show that litter quality strongly influences tadpole performance and suggest that functional traits, irrespective of species origin or identity, can have important consequences for ecosystem function.

Collins, Rachel J*; Bankert, Christina A; Gibson, Zach M; Patel, Devki G; Smith, Adrienne M – Roanoke College

Facilitation or apparent facilitation? Patterns of invasive plant and exotic earthworm abundances in Southwestern Virginia

Non-native species pose serious threats to native communities and are a top cause for species endangerment worldwide. Two important approaches to understanding these threats include assessing (1) the causes of invasion, and (2) the effect of invasions. We used both approaches in studying the effects of two invasive plant species (Asian Stilt grass (Microstegium vimineum) and garlic mustard (Alliaria petiolata)) and exotic earthworm communities in forest in Southwestern Virginia. We surveyed plant and earthworm communities in eight forests in 2007 and 2008. These sites ranged from small suburban fragments (<10 ha) to large nature reserves (5,100 ha). After two years of sampling, earthworm communities in our study sites appear to be comprised solely of exotic species. We found positive correlations between exotic earthworm richness and the abundances of exotic plant species and vines (multiple regression; r2 = 0.97 ; P < 0.5; n = 7 ). Further, exotic plant species richness was greater within areas of dense M. vimineum and A. petiolata populations compared to areas without invasive plant species (two sample t-test; t = 2.69; P < 0.05; df = 19). Lastly, we found a negative correlation between native plant species richness and invasive plant species abundance (linear regression; r2 = 18.3, P < 0.05, N = 24). Taken together, our results suggest that invasive plants and exotic earthworms may facilitate each other similar to the Invasion Meltdown mechanism and our sites seem to exhibit Biotic Resistance where high native species richness inhibits invasions.

Farrer, Emily C*; Goldberg, Deborah E – University of Michigan

Mechanisms and reversibility of effects of hybrid cattail on native wetlands

Most invasive plants negatively affect the native communities they invade. However, invasives can affect natives via different mechanisms, and distinguishing among them is essential for understanding the dynamics of invasions and for directing management strategies. Two common mechanisms of invasive plant effect are direct competition and alteration of the environment. We tested this in a Great Lakes marsh invaded by Typha x glauca (hybrid cattail), which produces monodominant stands with considerable litter accumulation. We teased apart the effects of live T. x glauca vs. its litter on the environment and plant community with a live/litter transplant experiment. We also tested whether these effects were reversible by a live/litter removal experiment in the Typha stand. Over four years, both transplanted litter and live T. x glauca increased soil inorganic nitrogen, but only litter decreased light. Litter had a much stronger effect on the plant community, decreasing native diversity and abundance and changing community composition to a more terrestrial species assemblage. Removal of litter and live T. x glauca had no affect on soil nutrients, but litter removal greatly increased light. Litter removal resulted in a greater increase in native diversity and abundance than removal of live T. x glauca, however it did not restore native marsh species, but rather recruited more terrestrial grasses and forbs. These results suggest that T. x glauca affects native plants by altering the environment by litter accumulation; however removal of aboveground litter will not restore wetland communities, because altered soil conditions favor colonization by terrestrial plants.

Giencke, Lisa M*; Dovčiak, Martin; Mitchell, Myron J – SUNY College of Environmental Science and Forestry

Spatial dynamics and effects of beech bark disease on an Adirondack forest

American beech (Fagus grandifolia), a major forest tree species in the Northeastern U.S., has been increasingly infected throughout its range by beech bark disease (BBD), which is caused by a non-native insect scale (Cryptococcus fagisuga) and fungal pathogens (non-native Neonectria coccinea var. faginata or native N. galligena). Although BBD can considerably affect forest ecosystem structure and function, our understanding of disease spatial dynamics in infected stands and its effects on tree recruitment is rudimentary. We studied spatial patterns of insect scale, canker and tree recruitment using historical stem maps from a ~2 ha plot in a mature northern hardwood forest in the central Adirondacks. Species, diameter at breast height (DBH) and spatial location of all trees (with DBH > 5 cm) in the plot were recorded in 1985 and 2000. Percentage of each beech stem infected by scale and canker was recorded in 2000. We analyzed spatial patterns of infected beech trees and tree recruitment using Ripley’s K-function. Almost 80% of small beech trees (DBH < 10 cm) were not infected by canker but about half were already attacked by scale in a spatially patchy fashion. The majority of larger beech trees were attacked by scale and canker; attack severity appeared correlated to distance from a heavily infested stand section. Despite increased mortality, beech abundance increased from 1985 to 2000; dead beech trees were positively associated with live beech recruits (but negatively with uncankered recruits). Thus, BBD can lead to continuous presence of low quality beech stands.

Jiang, Hao; Zimmerer, Edward; Liu, Dongjiao; He, Kate* – Murray State University

Using molecular markers to study the patterns of genotypic diversity of an invasive plant, alligator weed (Alternanthera philoxeroides) in southeastern US

Alligator weed (Alternanthera philoxeroides) is a successful invader native to South America. It has invaded all continents except Africa and Europe. In spite of its serious invasiveness all over the globe, alligator weed has been rarely studied in terms of its invasion mechanisms. It is even more surprising that knowledge on the relationships between its genetic variation and invasiveness is still very limited. This project attempts to uncover the mechanisms of alligator weed invasion using molecular markers to examine the patterns of genotypic variation of this successful invader. The analysis of genetic variation was carried out using Inter-Simple Sequence Repeat markers (ISSRs) on plant samples collected from three states in the southeastern US. The molecular evidence indicates that there is genetic variation in alligator weed populations. The results of this study suggest that genetic variation is closely related to the history of species introduction. Moreover, high genetic variation found in alligator weed populations contributes to its invasion success.

Jones, Chad C* – Connecticut College

Habitat distribution models poorly predict future occurrences of two invasive species

Habitat distribution models are increasingly used to predict the potential distribution of invasive species. If successful, these models can help managers target limited resources for monitoring and controlling invasive species to areas where the species are most likely to occur. The accuracy of these models is usually determined based on the current species distribution. However, because invasive species are not at equilibrium with the environment, models that accurately predict current distributions of invasive species may not be effective at predicting future distributions. I used data on the 1982 distributions of two invasive species, Celastrus orbiculatus and Rosa multiflora, in a natural area in southeastern Connecticut to create habitat distribution models using logistic regression. I then assessed how well these models predicted the distributions of the two species in 1982, 1992 and 2002. Overall model accuracy for Celastrus was high in 1982 and 1992 but declined in 2002. In addition, the ability of models to accurately predict new occurrences declined dramatically from 98% in 1982 to 76% in 1992 and only 27% in 2002. Models for Rosa followed a similar trend although predictive accuracy only dropped from 92% in 1982 to 61% in 2002. Both species have begun to invade types of habitats not invaded in 1982. These results suggest that measures of model accuracy from the time that model is developed are not good determinants of the ability of models to predict future occurrences. Thus habitat distribution models for invasive species must be used with great care.

Knight, Kathleen S*; Long, Robert P; Rebbeck, Joanne; Herms, Daniel A; Cardina, John; Herms, Catherine P; Klooster, Wendy; Gandhi, Kamal JK; Smith, Annemarie; Costilow, Kyle; Long, Lawrence; Smith, Stephanie K; Cappaert, David L – US Forest Service, Northern Research Station

Effects of emerald ash borer (Agrilus planipennis) on forest ecosystems in North America

Emerald ash borer (EAB) (Agrilus planipennis), a recently introduced beetle, has killed millions of ash (Fraxinus spp.) trees in North America. In over 250 monitoring plots in forests in Ohio and Michigan, representing a gradient of EAB infestation duration, we are monitoring the decline and mortality of thousands of ash trees, the responses of both native and invasive plant species, changes in species composition and forest structure, and effects on other organisms and ecosystem processes. The plots are located in forest stands of different ages and in different habitats to include all five ash tree species native to the region (F. Americana, F. pennsylvanica, F. nigra, F. profunda, and F. quadrangulata). Mortality of mature ash trees in a forest stand may reach 98% within six years of infestation by EAB, and is not affected by habitat or by the density, basal area, or species composition of ash trees. Invasive plant species are present, but not dominant, in most plots, with individual species distributions related to habitat, geography, and land use history. These plants may increase in abundance due to canopy gaps that result from ash mortality. EAB may dynamically interact with other species in these forest ecosystems. Woodpecker feeding behavior changes as woodpeckers take advantage of the abundant food source that EAB represents. Native parasitoid wasp species have been observed parasitizing EAB. We will continue to monitor these changing forest ecosystems as the effects of EAB reverberate through them.

Liu, Dongjiao*; Jiang, Hao; Zhang, Robin; He, Kate – Murray State University

Predicting the spatial distribution of an invasive plant, Lonicera japonica, based on species occurrence data from two watersheds in Western KY and TN

The invasion of alien plants has serious ecological and economic consequences. Geographic factors including human disturbances and habitat characteristics such as land cover, terrain, water and soils play an important role in plant invasion. However, the spatial distribution of most invasive plants is poorly documented, the path of dissemination is sketchy and the mechanism of spatial dispersal is mostly unclear. This project examines and compares the spatial distribution of a successful invasive plant, Japanese honeysuckle (Lonicera japonica) , in two watersheds of similar size but ecologically distinct in Western Kentucky (Ledbetter Creek) and Western Tennessee (Panther Creek). The occurrence data of Japanese honeysuckle and nine environmental variables were collected and measured from a total of 283 random plots at the two watersheds. A spatial logistic regression model was developed to identify the factors that contribute most to the spread of this invasive plant. Our results show that the spatial distribution of this invasive plant appears to be different at the two watersheds. The Ledbetter Creek watershed with heavier anthropogenic disturbances has a greater distribution of Japanese honeysuckle than the forested Panther Creek Watershed. The spatial regression model indicates that the distance from the main road, soil moisture, light intensity, and plant species richness of each plot were significantly correlated with the spatial distribution of invasive species at the Ledbetter Creek Watershed. However, elevation was the only significant factors in relation to the spatial distribution of Japanese honeysuckle at the Panther Creek Watershed. Furthermore, our results suggest that the invasion risk is strongly linked to anthropogenic disturbances.

Molofsky, Jane*; Collins, A Robin – University of Vermont

Evolution of range margins in invasive species

The study of invasive species at their current range margin and beyond can provide information on the likely distributional limits of the invasive species and its response to changing climate. In this study, we planted individuals of the invasive grass Phalaris arundinacea taken from its native range in Europe and its introduced range in the United States into common gardens along a climatic gradient to determine whether individuals are locally adapted to the current climate regime and whether they have the potential to evolve with changing climate. Native genotypes collected from the center (Czech Republic) and their range margin (France) and introduced genotypes collected from the center (Vermont) and the current range margin (North Carolina) were grown in identical common gardens in Vermont, North Carolina, Georgia and Florida. Survivorship declined sharply along this climatic gradient with no individuals surviving in Florida; there were no significant differences in survivorship between native and invasive genotypes. For tiller number and biomass, there were few significant differences among populations in their mean performance within a single garden; furthermore, there was no evidence for local adaptation of individuals to a particular climatic region. In North Carolina, both the French and North Carolina genotypes exhibited higher heritability for tiller number and biomass than the Czech Republic and Vermont genotypes. Thus, genotypes from climatically similar regions had higher evolutionary potential than those that came from climatically dissimilar regions. However, at the climatic extremes of Vermont and Georgia, we found that invasive North Carolina genotypes had significantly higher heritability than the native French genotypes.

Mooney, Emily H*; Morrissey, Danielle – Massachusetts College of Liberal Arts

Population variation in herbivory-related traits in Japanese knotweed, Fallopia japonica

Japanese knotweed, Fallopia japonica, is a non-native, invasive species throughout Europe and the United States. Our research focused on understanding how herbivory-related traits vary among populations of F. japonica. Plants were propagated from rhizomes obtained from three distinct populations. We used Spodoptera exigua larvae in a bioassay to assess variation among these population sources. Larvae were allowed to feed on plants for 72 hours, after which we measured the leaf area consumed by larvae and the weight gain by larvae. In addition, we assessed plants response to herbivory by measuring chlorophyll fluorescence. While weight gain by the larvae did not vary among population sources, there was evidence that response to herbivory varied among population sources. Specifically, chlorophyll fluorescence increased following herbivory, but the extent of increase varied among population sources. We are currently analyzing leaf area data, so that we can test for differences in conversion efficiency among larvae feeding on plants from different populations sources.

O'Donnell, Kelly L*; Pigliucci, Massimo – Stony Brook University

Selection dynamics of native and invasive annual Polygonum species

The ability to evolve rapidly is often implicated in discussions of what makes invasive plant species invasive. However, we are often lacking knowledge of which traits are under selection during an invasion. Plant invasions represent natural experiments that allow us to both explore the dynamics of natural selection in the wild and examine the evolution of an invader on contemporary time scales. We conducted a study of 10 natural populations of two invasive species (Persicaria lapathifolia and P. cespitosa) and one native species (P. pensylvanica) to quantify the amount of natural selection acting on these species to compare the selection dynamics to which each is exposed. We also conducted a germination trial to compare the potential for invasion determined by germination rate. A Lande-Arnold style multiple regression selection analysis was performed on five morphological traits (height, stem diameter, leaf number, leaf shape, and leaf area) using flower number as our fitness proxy. Most selection was indirect and caused by correlations with other traits under selection. However, there was significant direct selection for increased leaf number in both natives and invasives and thicker stems in just the invasives. The germination test showed that not only do the invasive plants have a significantly higher germination rate (>60% compared to 3%), but they also germinate significantly faster; both findings have large implications for the ability of these two invasives to spread. Another field season would need to be conducted to examine how each species is responding to their set of selection pressures.

Pinder, Rebecca A1*; Groffman, Peter M2; Robinson, George R1 – (1) University at Albany; (2) Cary Institute of Ecosystem Studies

Streamside interactions between salamanders, earthworms and soil nitrogen cycle processes

In North American forests, exotic earthworms are expanding their ranges into forests that were previously earthworm free. Earthworms can disrupt native forests by altering understory plant composition, consuming large amounts of leaf litter, mixing soil layers, and altering nutrient dynamics. The main objective of this study was to determine if quantity and diversity of earthworms and salamanders along stream banks are associated with differences in nitrogen cycling in these soils, focusing, in particular, on assessing the rates of nitrification, mineralization and denitrification along the stream bank. Soil samples were collected from stream banks in the Adirondack, Catskill and Helderberg regions of New York State where salamander and earthworm abundances, level of co-occurrence, and diversity were measured. Nitrification, mineralization and denitrification rates were compared across 16 streams. Potential denitrification rates were determined using a denitrification enzyme assay, while potential nitrification and mineralization rates were determined using laboratory incubations. There was a positive, though not significant, relationship between denitrification potential and the number of earthworms encountered. There was no significant relationship between nitrification and mineralization rates and abundances of earthworms or salamanders. As some earthworm species are thought to exert more of an effect on nitrogen cycle processes than others, further analysis will be done on a species by species basis (1237 earthworms were collected for identification and 1138 salamanders were identified to species in situ). Future work will focus on designing controlled experiments to further examine how earthworms and salamander interactions affect nitrogen cycling along the stream edge.

Rauschert, Emily*; Mortensen, David – The Pennsylvania State University

Quantifying the role of roads and road maintenance in the spread of an invasive plant, Microstegium vimineum

Preventing the spread of invasive species into new areas is seen as a critical for management, as eradication is often difficult, if not impossible, once populations are well established. The importance of human activity for the spread of invasive plants has long been recognized, but quantification of the impact of the vectors involved is lacking. We present a conceptual framework and ongoing experiments to understand the rapid spread of the invasive exotic Microstegium vimineum (Japanese stiltgrass). Detailed small-scale experiments provide the data to model the natural spread of Microstegium, which is several orders of magnitude slower than actual large-scale observations. Spread in roadside habitats, although variable, was found to be greater than in forested or wetland habitats. Forested roads, along which Microstegium grows abundantly, are often unpaved and are subject to frequent maintenance activities such as road grading. Experiments using seed proxies document the potential of road grading to rapidly spread seeds throughout road networks. We also investigate the role of road structures such as culverts in facilitating invasion into the forest interior. These results highlight the need to limit human-mediated spread when managing invasive species.

Rollinson, E*; Ness, JH – Skidmore College

Exotic plant species experience less herbivory than natives in a phylogenetically controlled survey

Enemy release is one mechanism by which exotic species may spread and become invasive. The enemy release hypothesis proposes that introduced plant species leave co-evolved natural enemies behind in their native ranges and, as a result, experience reduced herbivory and pathogen loads in their introduced range. This reduction in natural enemies may facilitate invasions of exotic plants if they experience lower levels of herbivory than native plant species. We compared herbivory on native and exotic plant species that naturally co-occur in riparian and forest habitats in Saratoga County, NY. We monitored a total of 43 plant species (26 native and 17 exotic; average 9 ± 4 individuals per species) in 30 families. Differences in herbivory between exotic and native species may be confounded by phylogenetic relationships among species in either of these categories; using taxonomically paired exotic and native species controls for these common ancestry effects. Therefore, we both directly compared native and exotic species, and compared them within taxonomically controlled groups. Exotic species had significantly lower levels of herbivory than did native species; this result remained when only taxonomically-controlled groups of species were compared. Our results suggest that observations of reduced herbivory in exotic species is truly a result of their introduced status, rather than an artifact of any common ancestry among exotic species. This enemy release may be an important mechanism of biological invasions.

Rollinson, E; Ness, JH* – Skidmore College

Consequences of riparian invasion for in-stream food webs and litter processing

Small streams are often powered by energy and other resources delivered by senescing leaves falling from riparian vegetation. We tested the hypothesis that invasions of riparian zones by Japanese knotweed (Fallopia japonica) alter the development of in-stream food webs. We used 0.1m2 buckets to collect dehiscing leaves along a transect that included invaded and non-invaded riparian habitat abutting Kayaderosseras Creek, NY. We weighed the leaves that fell weekly from 21 Sept to 25 Nov, 2008. Owing to among-species differences in leaf-fall phenology (median loss prior to Oct 14 for Populus, Acer, Rhamnus, Tilia and Ulmus spp, Oct 28 for Quercus spp and Nov 4 for Fallopia), leaves in knotweed-invaded microsites entered streams later in the autumn. We created leaf-packs that mimicked the changing composition of in-stream litter derived from these two habitats, systematically adding leaf material to leaf packs and collecting a sub-sample of the leaf-packs five times over a 40 day interval. We demonstrated that litter from invaded microsites was processed more slowly over a given time interval and also found differences in macro-invertebrate community composition, with higher diversity and lower Plecoptera representation in the invaded leaf-packs. We attribute these differences to the combined effects of differences in leaf quality, leaf age, and progressively falling water temperatures when late autumn leaves, such as knotweed, enter the stream.

Wildova, Radka*123; Goldberg, Deborah1 – (1) University of Michigan; (2) Institute of Botany, Academy of Sciences of the Czech Republic; (3) Ecological Research Institute

Exotic and native cattails and their hybrid in North America: Implications for understanding cattail invasions

We examined the ecology of invasions of northern Michigan wetlands by the European cattail, Typha angustifolia, and its hybrid, T. x glauca, with native T. latifolia. Little is known about wetland characteristics affecting invasibility, the mechanisms by which T. angustifolia or T. x glauca invades, or whether either displaces T. latifolia. Surveys revealed T. angustifolia was more abundant in unshaded, densely-vegetated wetlands on sandy soils, while T. latifolia was better represented at sites with exposed organic soils, standing water, and tree shading. The hybrid was restricted to unshaded sites with fine textured soils. However, all three parent taxa did have some habitat overlap. Two experiments revealed responses to environmental conditions consistent with these patterns. In a common garden experiment, low soil organic matter (SOM) decreased the growth of T. latifolia and the hybrid, but not that of T. angustifolia. In contrast, low light decreased the growth of T. angustifolia and the hybrid, but not of T. latifolia. The hybrid had higher growth than both parents only when both SOM and light were high. However, in a four year field experiment in which all three taxa were transplanted to a marsh with sandy soil, low SOM and high light, both survival and productivity of the hybrid were consistently higher than both parents despite these oligotrophic conditions. Our results suggest differing growth responses to soil texture, organic matter, and light can enable invasion by both the non-native and hybrid taxa into wetland habitats that would be unoccupied by the native species, and that degree of eutrophication is not the key variable triggering invasion.

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